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Totoro is a forest spirit that little Mei, and later her older sister Satsuki, encounter in a giant camphor tree near their new home in the countryside. Although their father, a university professor, is with them when they move, their mother is in the hospital, recovering from some unnamed illness.

The story centers on individuals who are gifted with super natural powers and use them for different purposes including holding a business, solving mysteries, and carrying out missions assigned by the mafia. The story mainly follows the of the "Armed Detective Agency" and their everyday lives.